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The researchers evaluated the accuracy of their maps by comparing them with other existing topographic models, looking for discrepancies or errors in lunar surface features.

Atmospheric simulations, for example, must account for complex variables such as temperature changes and air turbulence under different topographic conditions.

These similarities are despite any differences in glacial history, background topographic steepness, annual precipitation, and other factors that would otherwise govern the push and pull of water and land, the researchers report.

You can clearly see these on a topographic map as circles or ovals that have small lines pointing to the center of the circle or oval.

In short, the traits of each species correspond to fitness peaks that can be likened to mountains on a topographic map separated from other mountains by valleys of lower fitness.